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As a biologist trained in taxonomy, Josie's primary task was studying and classifying plants and animals on the moon. In Red Sun Rising, 236 years ago, Josephine and her family were among the original families (the Primes) to settle on Sanctum as part of Mission Team Alpha. Josephine was on her seventh host, making her Josephine VIII. Her family became three of the Primes and used various hosts for the next two centuries to be reborn after each death. Gabriel, who had terminal cancer by the time Josephine was resurrected, died soon afterwards and Josephine brought him back as she couldn't bear to lose Gabriel. Josephine was the first person to be brought back using this technology. In 2070, after 25 years of failed experiments, Gabriel was able to figure out how to use the Mind Drives to bring their loved ones back by uploading their entire consciousness into a host with black blood. The pain of losing Josephine pushed her father and her boyfriend, Gabriel Santiago, to start exploring ways to bring her back. Josephine was killed by her father, who was suffering from the psychosis.

Unknown to them, the eclipse triggers the release of a psychosis-inducing toxin from the plants. On day 21 of the mission, the team endured an eclipse of the two suns. It was Josephine who suggested naming the place 'Sanctum' - the Latin-root for sanctuary. She was in charge of classifying lifeforms on their new world. Josephine was one of the biologists for the team. They arrived there in 2045 and were original members of Mission Team Alpha. Josephine and her parents went aboard Eligius III to travel to the moon Alpha. She originally was not going to go with her parents on the Mission, but this event caused her to go.

In December 2043, she and her friend Olivia were in a diner in Boston, Massachusetts, USA, when her stalker Dave came in and shot himself in the head because she ignored him. Josephine was born on Earth to Simone and Russell Lightbourne.

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